How did brachiopods go extinct.

How did brachiopods go extinct Widespread families of trilobites disappeared and graptolites came close to total extinction. Groups of trilobites disappeared at each of the three extinctions and very few survived into the following Carboniferous Period. Ash from southwestern China’s Emeishan Traps, for example, dates to the Capitanian and has previously been implicated as a potential cause of the local brachiopod extinction. In some genera (e. Here, new Bayesian analyses suggest May 6, 2014 · Among the carcasses are dozens of species of small shelled marine organisms called brachiopods, their tight-lipped expressions frozen in time. Within the marine paleoenvironment, trilobites were found in a broad range from extremely shallow water to very deep water. This group survived the Great Permian Extinction and eventually became extinct during the Jurassic Period. Brachiopods in some ways resemble clams but differ from clams in shell symmetry. Bivalves –– 1. V geological sciences, is using these brachiopods to investigate patterns of extinction and species formation over time, including during several of the major extinction events that have happened on Brachiopods are marine invertebrate animals with two shells. Can extinct animals be revived? Scientists have made some progress in reviving extinct species through cloning and genetic engineering, but such attempts have had limited Study with Quizlet and memorize flashcards containing terms like Terrestrial organisms such as dinosaurs are far less likely to be fossilized than marine organisms such as brachiopods or corals. Late Devonian extinction - 383-359 million years ago Sep 1, 2016 · The orders Rhynchonellida and Terebratulida diversified in the Jurassic and are still extant. Classification: Phylum: Brachiopoda ; Class: Articulata; Order: Spiriferida; Genus What major events happened during the Silurian period? Possibly the most remarkable biological event during the Silurian was the evolution and diversification of fish. Bivalves go extinct at the P-T mass extinction, brachiopods do not . Mar 13, 2025 · As the article states, “Marine reptiles, such as ichthyosaurs, plesiosaurs and mosasaurs are not dinosaurs… None of these other extinct groups shared the characteristic upright stance of dinosaurs. When did plesiosaurs go extinct? Plesiosaurs went extinct approximately 66 million years ago during the K-Pg extinction event. Did all sponges go extinct at the end of the Triassic period? No, while some species of sponges went extinct at the end of the Triassic period, the group as a whole did Jul 12, 2024 · In fact, Monarrez notes, the baseline rate at which species normally go extinct has decreased over time. • The Terebratulida don’t make their first appearance until the Carboniferous Period. Mucrospirifer ) it is greatly elongated, giving them a wing-like appearance. Oct 7, 2024 · According to Claus Nielsen’s (1991) ‘brachiopod fold’ hypothesis (later adapted by Cohen and colleagues in 2003), brachiopods are descendants of an ancestor similar to Halkieria, a small, slug-like animal. The “Ordovician radiation” which followed the late Cambrian extinctions, lead to a tripling of marine diversity, the greatest increase in the history of life, and giving the highest levels of diversity seen during the Paleozoic Era. Spiriferids and brachiopods in general, hit the height of diversity during the Devonian Period. Jul 5, 2023 · Trilobites are a group of extinct marine arthropods that existed for a staggering period of time, spanning over 270 million years. Oct 25, 2024 · Brachiopods, sometimes called “lamp shells,” filled many of the ecological niches in Paleozoic oceans that bivalves have occupied in Mesozoic and Cenozoic oceans after approximately 95% of brachiopods species became extinct at the end of the Paleozoic. 5. The internal organs and muscular systems of clams Although many rhynchonelliform brachiopods are held in place by a pedicle, some extinct forms lost the pedicle and lay freely on the sea bottom. . e. … Before the extinction event, brachiopods were more numerous and diverse than bivalve mollusks. Oct 25, 2024 · Brachiopods, sometimes called “lamp shells,” filled many of the ecological niches in Paleozoic oceans that bivalves have occupied in Mesozoic and Cenozoic oceans after approximately 95% of brachiopods species became extinct at the end of the Paleozoic. Lamp shells - Fossilization, Mollusks, Brachiopods: Brachiopods were among the first animals to appear at the beginning of the Cambrian Period. A millipede-like organism is inferred because the burrows occur in discrete size classes, are bilaterally symmetrical, and were backfilled Brachiopods (Figure 7. Most brachiopods are attached to the substrate by means of a fleshy "stalk" or pedicle. Other reptile-like mammal groups continued to do well in the Permian including the anthracosaurs and seymouriamorphs. Concavo-convex; Radiating striae which alternate in size. Trilobites are one of the earliest known complex life forms and left Sep 9, 2023 · However, brachiopod orders that survived or originated after the PTME (PTs brachiopods, g–i) did not show an apparent decline in diversity in Triassic–Jurassic. Jan 11, 2022 · Andrew Bush of the University of Connecticut, the paper's senior author, says it was surprising to discover that the first pulse was more detrimental to the brachiopods. 4 Brachiopod PreservationAbove Image: Animal forms; a second book of zoology (1902), Figure 43: Animals of Uncertain Relationships. 1 Brachiopod Classification –– 1. Members from the orders Lingulata, Rhynconellida, and Terebratulida are among those that exist today. , a variety of shell shapes) over time. eScholarship It has been suggested that the slow decline of the brachiopods over the last 100 million years or so is a direct result of the rise in diversity of filter-feeding bivalves, which have ousted the brachiopods from their former habitats; however, the bivalves have undergone a steady rise in diversity from the mid-Paleozoic onwards, and their Sep 14, 2023 · One of the biggest crises in Earth history was marked by a revolution in the shellfish – brachiopods, sometimes called ‘lamp shells’ were replaced everywhere ecologically by the bivalves, such as oysters and clams. The hypothesis proposes that the first brachiopod folded the rear part of its body under its front, giving rise to the paired valves. The greatest recorded mass extinction in Earth’s history occurred at the end of the Permian Period. Mar 27, 2019 · The order eventually went extinct in a subsequent mass extinction, the scientists said. These are the babies of the group a mere 350 million years old. When did they live? The oldest brachiopods can be found in rocks of early Cambrian age (about 530 million years old). " Brachiopod shells may be either phosphatic or (in most groups) calcitic. The Permian-Triassic mass extinction further contracted the total occupied morphospace, dividing the survivors into two distinct groups, with the linguloids and discinoids exclusively found in the areas of morphospace represented by low Aug 10, 2012 · The Ordovician* lasted about 45 million years and saw the transition from very primitive to relatively modern life-forms in the seas. While Brachiosaurus was a terrestrial dinosaur, the cascading effects of such anoxic events on Brachiopods associated with reefs also became extinct. It was also noted that analyses of ‘homogeneous sites only’ and ‘heterogeneous sites only’ datasets resulted in different Mar 10, 2025 · Archaeocyathans were an ancient group of sponges that went extinct, but they left no living descendants. The acquisition of new, and the redescription of existing faunas, in precise spatial and temporal frameworks, using new and well‐established analytical and investigative techniques, are ¨ Mass extinctions occur when anomalously high numbers of species go extinct in a short period of time. , Fossils are _____. Brachiopods were more abundant in the Paleozoic, bivalves were more abundant in the Mesozoic . Brachiopoda were a dominant group of marine organisms during the Brachiopods (/ ˈ b r æ k i oʊ ˌ p ɒ d /), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. N O V, D. Sep 9, 2023 · Brachiopod-bivalve switch in diversity dominance after the Palaeozoic era is a textbook example of clade replacement, and its mechanism has long been debated. What are the 5 extinctions? Top Five Extinctions Study with Quizlet and memorize flashcards containing terms like what are brachiopods?, when did the brachiopod originate?, when did many brachiopods go extinct? and more. Today this is the most abundant of the brachiopods. The range chart of the orders of Rhyn- Trilobites (/ ˈ t r aɪ l ə ˌ b aɪ t s, ˈ t r ɪ l ə-/; [4] [5] [6] meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. Values above 0. Who became extinct? All of the major animal groups of the Ordovician oceans survived, including trilobites, brachiopods, corals, crinoids and graptolites, but each lost important members. Examination of organismal characteristics which promote survivorship through both background and mass extinctions may reveal general ecological principles potentially critical to modern conservation efforts. Aug 1, 2007 · From these researches, it has been well documented that four major brachiopod orders (Productida, Spiriferida, Orthida, Orthotetida) became extinct near the Permian-Triassic boundary; while four Since some 95% of all brachiopod taxa are extinct, the fossil record is the primary source of data to frame and test models for the evolution of the phylum. They might just look like clams, but they are not even closely related. They possess a lophophore, excretory organs (nephridia), and simple circulatory, nervous, and reproductive systems. … Apr 27, 2016 · Bottom: partial dependence plots for GBM models trained on extinction patterns in each interval. Straight hinge line; Bilobed cardinal process; Ventral valve does not have a strongly limited muscle area, Oct 20, 2023 · In contrast, while the Ordovician-Silurian extinction event was significant, it did not specifically target brachiopods to the same extent as seen during the late Devonian extinction. Brachiopods open when they die, bivalves Sep 20, 2019 · became extinct at the Brachiopod Big Five (indicated by ar-rows). , Dec. 10. They first appeared during the Early Cambrian period around 521 million years ago and became extinct during the mass extinction event at the end of the Permian period around 252 million years ago. 1. Examples of groups of brachiopods and trilobites that became extinct are shown below. Although Brachiopod larvae swim about freely, the adults are frequently anchored or cemented to objects on the sea floor by a fleshy stalke (pedicle) or by spines. Study with Quizlet and memorize flashcards containing terms like brachiopoda, pedicle, pedicle/ventral valve and more. Brachiopod shells come in a variety of shapes and sizes. Aug 20, 2007 · Brachiopod faunas were very abundant and diversified in the marine realm during the Late Paleozoic, but were drastically reduced in species richness in the Early Triassic after nearly 87–90% of genera and 94–96% of species became extinct at the end of the Permian (Shi and Shen, 2000, Shen and Shi, 2002). 3. Estimating the Time Separating Extinction Pulses: A Graphical Procedure Because we have rejected a simultaneous extinction time for ostracodes and brachiopods, it is natural to ask how much time separated their two extinction pulses ( Fig. Sclafani took existing evolutionary tree information for the brachiopods and used computer models to analyze changes in shell structure among dozens of different groups in the order and to place those changes on a timeline. Jun 27, 2017 · Since some 95% of all brachiopod taxa are extinct, the fossil record is the primary source of data to frame and test models for the evolution of the phylum. B. Jul 7, 2022 · How did brachiopods go extinct? Besides marking the disappearance of species, the Capitanian was also a time of major volcanic eruptions . The extinction appears to have occurred in several phases. The Ordovician Period was a time of significant geological, climatic, and biological changes. 2A) supported brachiopods and phoronids as sister groups, with the monophyly of brachiopods moderately supported (posterior probability of 0. Finer growth lines crossing the radiating striae. Since some 95% of all brachiopod taxa are extinct, the fossil Dec 22, 2007 · I did some reading and found some theories on why some think most of the abundant brachiopods died off (95% of species) while the pelecypods prospered so well. Although their body was strongly built, storms would occasionally destroy the stromatoporoid reefs. Modern day brachiopods do still exist in the form of lingula. Phylum Brachiopoda. In summary, the correct answer to the question of when more than half of brachiopod species became extinct is at the end of the Devonian Period. Which of the following statements best explains this discrepancy between terrestrial and marine animal fossilization?, Evidence of an once-living organism preserved in sedimentary rock is an example of Permian–Triassic boundary at Frazer Beach in New South Wales, with the End Permian extinction event located just above the coal layer [2]. g. Jul 7, 2022 · Brachiopods are extremely common fossils throughout the Palaeozoic. Brachiopods and bivalves both permanently attach to the seafloor . Mar 8, 2025 · 9. Which of the following statements best explains this discrepancy between terrestrial and marine animal fossilization? Jul 15, 2022 · Trilobites appear to have been exclusively marine organisms, since the fossilized remains of trilobites are always found in rocks containing fossils of other salt-water animals such as brachiopods, crinoids and corals. Is Brachiopod a species? Mucrospirifer, genus of extinct brachiopods (lamp shells) found as fossils in Middle and Upper Devonian marine rocks (the Devonian Period began 416 million years ago and lasted about 57 million years). Feb 28, 2025 · It is mostly seen in the fossil record of marine invertebrates: many brachiopods, trilobites, bryozoans, and graptolites became extinct in two short pulses separated by a geologically short time. Mar 11, 2025 · The earliest known mass extinction occurred during the Ordovician period. It was also noted that analyses of ‘homogeneous sites only’ and ‘heterogeneous sites only’ datasets resulted in different Rafinesquina is an extinct genus of facultatively mobile, epifaunal suspension feeders. Brachiopods are also particularly suitable for palaeoecological analyses. Only about 300 to 500 species of brachiopods exist today, a small fraction of the perhaps 15,000 species (living and extinct) that make up the phylum Brachiopoda. 4. At left is a Rhynchonellida brachiopod. "We have to compare the samples before and after to get a sense of what survives and what completely disappears and goes extinct," Bush says. Jul 13, 2015 · The biggest of these happened toward the end of the Permian Period about 252 million years ago, when 95 percent of all species went extinct. Some paleontologists suggest that an early phase affecting graptolites, brachiopods, and trilobites took place prior to the end of the Ordovician Period, before the major fall in sea level occurred, and it may have been caused by falling carbon dioxide levels associated with the erosion of silicate rocks, which may have triggered a Western Australian Museum | Western Australian Museum Brachiopods. How might humans go extinct? Multiple threats could lead to human extinction, including nuclear war, climate change, biological warfare, and ecological collapse. Figure 11. Can you eat brachiopods? Aug 15, 2022 · Why did brachiopods go extinct? Anoxia would have resulted from a rise in temperature caused by elevated levels of heat-trapping carbon dioxide in the atmosphere, as oxygen doesn’t dissolve as well in warm water. However they did not become abundant until the Devonian. Jun 30, 2016 · thecideides were paedomorphic descendants of one group of extinct, spire-bearing brachiopods (spiriferides), rather than strophomenates. 5 indicate a tendency for genera with the given predictor value to go extinct when all other variables are held constant, values below 0. C. Terrestrial organisms such as dinosaurs are far less likely to be fossilized than marine organisms such as brachiopods or corals. Nov 14, 2022 · Most brachiopods became extinct about 250 million years ago during the P-T Extinction period. 2. The surviving brachiopods were mainly Productida, followed by Spiriferida. that did not go extinct underwent significant morphological changes during the gap between Permian ancestors and their Triassic descen- dants, which could reflect either opportunistic evolution after the extinction, or adaptive changes necessitated by environmental con- ditions. Instead of being horizontally symmetrical along their hinge, like clams and other bivalves, they are vertically symmetrical, cut down the middle of their shell. Overall, about 86% of species, 57% of genera, and 27% of taxonomic families died out, making this the second largest extinction in the Phanerozoic. Afterwards, in the Mesozoic, their diversity and numbers were drastically reduced and they were largely replaced by bivalve molluscs . Phylum Brachiopoda (lamp shells) has about 300 living species placed into two classes, Articulata and Inarticulata. 4 ). This changed after the mass extinction at the end Jun 27, 2017 · 2011, fig. Study with Quizlet and memorize flashcards containing terms like when did brachiopods appear in the fossil record and when did they reach their peak diversity? Which mass extinction impacted them severely? Are they extinct?, How do brachiopod shells grow? How does this contrast with trilobites?, What is the main mode of life of brachiopods? How do they feed? and more. [1] Reptile-like amphibians would go extinct in the Early Triassic, but one of their evolutionary lines, the amniote clade, would live on. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. Brachiopods are an ancient group of organisms, at least 600 million years old. 5 indicate a tendency for genera with the given predictor value to survive. Not only does this time period mark the wide and rapid spread of jawless fish but also the appearances of both the first known freshwater fish and the first fish with jaws. Why did brachiopods go extinct Aug 20, 2007 · Brachiopod faunas were very abundant and diversified in the marine realm during the Late Paleozoic, but were drastically reduced in species richness in the Early Triassic after nearly 87–90% of genera and 94–96% of species became extinct at the end of the Permian (Shi and Shen, 2000, Shen and Shi, 2002). In recent years, a number of species from the remote Ulutau mountains region of Kazakhstan have invaded the world market, with many of these dolomitic specimens (including Ditomopyge kumpani and Griffithides praepermicus) being fossilized along-side other fauna, including brachiopods and crinoids, providing an interesting view of life at this Brachiopod structure seems to have evolved in a series of steps: first a stationary filter feeder with a tubular shell (such as Eccentrotheca, a basal tommotiid brachiopod), second a bivalved shell which did not completely enclose the body (most tommotiids), and finally a bivalved shell which completely enclosed the body. Brachiopod morphology and terminology; Brachiopods versus bivalves Brachiopods superficially resemble clams but are not closely related to our modern sea shells. (Courtesy of the University of Ottowa) Between 100 and 10,000 of them go extinct They were mainly in shallow waters but could be found up to 110 meters. Oct 25, 2019 · Only 5% of all brachiopod species to ever exist still survive today, while 95% have gone extinct. A. "There are always species going extinct and new Why did the Brachiopod go extinct? Anoxia would have resulted from a rise in temperature caused by elevated levels of heat-trapping carbon dioxide in the atmosphere, as oxygen doesn’t dissolve as well in warm water. Mar 1, 2007 · We therefore reject H 0 and conclude that ostracodes and brachiopods did not go extinct at the same time. Study with Quizlet and memorize flashcards containing terms like A(n) ____ extinction involves the massive die-off of many millions of different ____ of organisms in a geologically short period of time. More than 30,000 Jul 10, 2023 · Which of these is true regarding brachiopods and bivalves? A. 3 Brachiopod Paleoecology ← –– 1. Mass extinctions and clade extinctions in the history of brachiopods 713. Spiriferida is an order of extinct articulate brachiopod fossils which are known for their long hinge-line, which is often the widest part of the shell. 9) range from the Lower Cambrian to the present. Compared to hundreds of species Sep 1, 2016 · The orders Rhynchonellida and Terebratulida diversified in the Jurassic and are still extant. 7 Ma after the end-Permian mass extinction. Major casualties were marine invertebrates like brachiopods, trilobites, bivalves, and corals. , Two requirements are usually necessary for an organism to become fossilized: _____. Sep 12, 2022 · Why did brachiopods go extinct? Anoxia would have resulted from a rise in temperature caused by elevated levels of heat-trapping carbon dioxide in the atmosphere, as oxygen doesn’t dissolve as well in warm water. ogy and life modes of the first brachiopods; (2) under-standing the relationships of the major groups to each other and higher sister taxa; and (3) unravelling the roles of the Cambrian and Ordovician radiations that set the agenda for much of subsequent brachiopod evolution. [109] Chapter contents: 1. Brachiopoda –– 1. Sep 26, 2019 · The event took its hardest toll on marine organisms such as corals, shelled brachiopods, eel-like creatures called conodonts, and the trilobites. Rarely Jul 7, 2022 · How do brachiopods eat? How did brachiopods go extinct? and including many extinct species commonly found as fossils. , B O L GO V A, G. They were at peak diversity in the Devonian, but most went extinct at the end of the Permian. The post-extinction brachiopods were also affected by a subsequent crisis corresponding to the boundary between MFB 2 and MFB 3 so that most survivors were extinct approximately 0. The Extinct Orders • Oblellida-was present and became extinct during the Cambrian Period. During the Paleozoic era (542-250 million years ago), brachiopods were one of the most abundant and diverse groups of marine organisms. 2 Brachiopods vs. Most people are not familiar with living brachiopods because modern species inhabit extremely deep regions of the world’s oceans, and their shells are rarely found on modern seashores. 23, 2024 These Brachiopod fossils date back to the time of the Late Ordovician mass extinction. But during the Paleozoic, thousands of different species of brachiopods teemed in the near-shore and deep-sea environments of Wisconsin. Modern lingulate brachiopods burrow into sand and mud on the sea floor. The best record across the They are currently the most successful and diverse group of living reptiles, with more than 10,000 extant species. In a new study, scientists have proposed that a sixth global extinction, about 10 million years before the End-Permian die-off, should be added to the list. On the other hand, the other two orders, Athyridida and Spiriferinida, were severely decimated by the end-Triassic crisis and became extinct in the Early Jurassic, during the second-order Toarcian extinction event, coincident with the Toarcian Oceanic Anoxic Event (T-OAE). It may be that life has survived so much since its origins that new species have become What is palæontology? Literally, the word translates from Greek παλαιός + ον + λόγος [ old + being + science ] and is the science that unravels the æons-long story of life on the planet Earth, from the earliest monera to the endless forms we have now, including humans, and of the various long-dead offshoots that still inspire today. Below are a few examples of some of these living brachiopods, which will be explained in more detail on the next page. Stromatoporoids were often found in the same strata with gastropods, articulate brachiopods, and rugose and tabulate corals (Figure 3). Sometimes the bottom valve is convex like the top valve, but in many species the bottom valve is concave or occasionally conical. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. In contrast, although some bivalves (such as oysters, mussels, and the extinct rudists) are fixed to the substrate, most are free-moving, usually by means of a muscular "foot. These events, characterized by widespread depletion of oxygen in the oceans, have been linked to mass extinctions in marine environments. I read that pelecypods use an energetically-efficient ligament-muscle system for opening valves, and thus require less food to subsist. The only major group of terrestrial lizards to go extinct at the end of the Cretaceous were the polyglyphanodontians, a diverse group of mainly herbivorous lizards known predominantly from the Northern Hemisphere. Influenced by such factors as water depth, salinity, oxygen levels and static lifestyle, the distribution patterns of fossil brachiopods provide a useful tool in deducing the position of ancient shorelines and the past distribution of land and sea. Source: Wikimedia Commons (Public Domain). The Early Toarcian anoxic event, for instance, severely impacted brachiopod populations and other marine fauna 2. Characteristics of the Genus. 82) and placed craniiforms at the base of the brachiopod clade. On average, every 4 million years a species will go extinct. Brachiopod fossils are often well-preserved, as well as being abundant and exhibiting diverse shell morphology (i. ” 2. It’s impossible to pinpoint a single “first” animal to go extinct, as extinction is a continuous process. How did the dodo bird go extinct? Apr 6, 2025 · Ordovician Period - Marine Life, Trilobites, Brachiopods: Although no fossils of land animals are known from the Ordovician, burrows and trackways from the Late Ordovician of Pennsylvania have been interpreted as produced by animals similar to millipedes. Brachiopods, which need oxygen, could have succumbed under such conditions. Mucrospirifer forms are characterized by an extended hinge line of the two valves, or shells, of 1936 Thylacine (Tasmanian tiger or wolf) – extinct from hunting, habitat loss, and competition with dogs; 1952 Deepwater cisco fish – extinct from competition and predation by introduced fishes; 1962 Hawaii chaff flower – extinct from habitat conversion to military installations; 1989 Golden toad – extinct from climate change or other Brachiopods are the most abundant fossils in Wisconsin. and more. Clams, or bivalves, belong to the Class Bivalvia in the Phylum Mollusca, while brachiopods belong to their own phylum, Brachiopoda. Brachiopoda were a dominant group of marine organisms during the Jul 9, 2022 · Mucrospirifer, genus of extinct brachiopods (lamp shells) found as fossils in Middle and Upper Devonian marine rocks (the Devonian Period began 416 million years ago and lasted about 57 million years). This study explores survivorship of brachiopods, a highly diverse and abundant Paleozoic clade, through the mid-Permian to mid-Triassic interval, which includes the greatest mass The Late Ordovician mass extinction (LOME), sometimes known as the end-Ordovician mass extinction or the Ordovician-Silurian extinction, is the first of the "big five" major mass extinction events in Earth's history, occurring roughly 445 million years ago (Ma). Overview Brachiopods are solitary creatures that inhabit the seafloor Dec 23, 2024 · Mon. 6: The most common fossils in Ordovician rocks are the brachiopods. This happened as a result of the devastating end-Permian mass extinction which reset the evolution of life 250 million years ago. The Permian–Triassic extinction event (also known as the P–T extinction event, the Late Permian extinction event, [3] the Latest Permian extinction event, [4] the End-Permian extinction event, [5] [6] and colloquially as the Great Dying, [7] [8]) was Apr 24, 2023 · The acrotheloids go extinct at the end Ordovician mass extinction, while the discinoids survive (Figure 2B). D. They were significant reef builders during the early Cambrian period. Although they outwardly resemble clams (which are bivalve mollusks), they are not closely related and their internal anatomy is completely different. In meticulous investigations of morphology and on- Jun 27, 2017 · 2011, fig. afux grhzq aimn idfd oozmxo ipmhv ldyj fkxvyhn ycvz fvbamea qcw ltuo liwexly ajzkv kxfxps